for natural selection will always act according to the nature of the places which are either unoccupied or not perfectly occupied by other beings
 Charles Darwin, On The Origin of Species (1859). copy citation

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Author Charles Darwin
Source On The Origin of Species
Topic selection acts
Date 1859
Language English
Reference
Note
Weblink http://www.gutenberg.org/files/2009/2009-h/2009-h.htm

Context

“it is far more probable that each form remains for long periods unaltered, and then again undergoes modification. Nor do I suppose that the most divergent varieties are invariably preserved: a medium form may often long endure, and may or may not produce more than one modified descendant; for natural selection will always act according to the nature of the places which are either unoccupied or not perfectly occupied by other beings; and this will depend on infinitely complex relations. But as a general rule, the more diversified in structure the descendants from any one species can be rendered, the more places they will be enabled to seize on, and the more their modified progeny will increase.” source