John M. Legler, A New Tortoise, Genus Gopherus…
“ Femoral spurs are developed to varying degrees in the other species of Gopherus; they are usually lacking in individuals of small to medium size. In large specimens of agassizii and polyphemus that I have examined, there is but one enlarged spur, although several pointed osteodermal scales may be present on the back of the thigh. ”
