by William Swainson, Zoological Illustrations — Vol II. Pl… (1820)
“ They are subject to variation in their colour; and this, joined with a general resemblance in form, has rendered the discrimination of the species very difficult. I have, however, remarked, that the various modifications of the umbilicus, and the termination of the pillar (which is indicated in many species by an elevated ridge or rib within the umbilicus) is a certain and constant indication, presenting the same peculiarity through all the individuals of a species, even in the young state. ”
