William Edward Duellman, Descriptions of Two Species of Frogs…
“ Stuart (1954:169) discussed the generic characters and pointed out that both the ventrolateral glands and horny nuptial spines were seasonal in their development, being found only in breeding males. Stuart then went on to describe Ptychohyla schmidtorum, a species characterized by the absence of horny nuptial spines in breeding males. My investigations of these frogs have revealed the presence of two groups of species. In both groups breeding males have large ventrolateral glands, but the two groups are easily separated by four characters. ”
