George Vernon Hudson,
An Elementary Manual of New Zealand Entomology…
(1892)
“ Although so very different in general appearance to the preceding insects, this beetle will be found on careful examination to agree with them in all important respects, being only what a ground beetle might naturally become if forced to lead an aquatic existence. Breathing is effected in all the water beetles by the spiracles of the abdomen, which alone are developed. The air is taken in between the elytra and the body, and owing to the convexity of the former, a supply can be retained sufficient to last the insect some twenty or thirty minutes. ”
