William Henry Flower and Henry Scherren, 1911 Encyclopædia Britannica (1911)
“ In the fore feet the web not only fills the inter spaces between the toes, but extends considerably beyond the ends of the long, broad and somewhat flattened nails, giving great expanse to the foot when used for swimming, though capable of being folded back on the palm when the animal is burrowing or walking on the land. On the hind foot the nails are long, curved and pointed, and the web extends only to their base. On the heel of the male is a strong, curved sharply pointed, movable horny spur, directed upwards and backwards, attached by its expanded base to the accessory bone of the tarsus. ”
