Erik Norman Kjellesvig-Waering, Eurypterids of the Devonian Holland Quarry Shale of Ohio (1961)
“ In contrast to these active swimmers, the associated eurypterids, namely, Dolichopterus asperatus and Syntomopterus richardsoni, had simple, small chelicerae not serviceable as weapons of either defense or predation; both were probably benthonic forms, crawling on the mud but perhaps capable of swimming to a limited extent. These could well have been prey to the fishes and, indeed, to the large pterygotids. However, in this shale the fishes are the only animals with a covering rigid and hard enough to abrade the tough pincer-teeth of the pterygotids. ”
