Bruce Fink, Ohio Biological Survey, Bull. 10…
“ Within the Lecideaceae, the line of evolution seems to have been in the direction of a well-developed exciple and from simpler to more complex spores. With the advance in these two directions has gone a slightly increased development of the thallus.In structure, the thallus is crustose, and the thalli vary from inconspicuous, evanescent conditions to those which are conspicuous and sometimes even subsquamulous. Rarely the thallus extends upward as a veil which surrounds the apothecia laterally and suggests how the thalloid exciple of higher families probably arose. ”
