T. A. Coward

“ The fact that birds fly in any direction in a wind, and when at low elevation pay little heed to the direction of the wind, when the breeze is light, simply means that they can fly faster than the medium they are in; if the medium travels faster than they do, they will be carried in it to their advantage or disadvantage.
Even in these days of upper-air investigation we really know very little about the. speed, direction and steadiness of upper-air currents; but we do know that at a moderate elevation—some two or three thousand feet—the strength is usually greater than nearer the earth. ”

“ The migrating bird, when passing between the breeding home and the winter quarters, travels by what is termed its Route. The definition of the route has caused more controversy than perhaps any other incident of migration; the chief point at issue is whether the bird uses a particular high road, along which all its fellows from the same area travel, or if all birds move in what has been called a "Broad Front." Ornithologists have been, and to some extent still are, divided into two camps, one upholding defined routes and the other the extended or broad front movement. ”

“ Sight, however, cannot be always necessary, for, at the Kentish Knock Lightship, Mr Eagle Clarke noticed that birds flew so low along the water that they could not possibly have seen their way.
It is purely speculative to say that the young bird which travels alone, for it seems certain that many young do travel unguided on their first journey, has an inherited memory of the actual route to be followed, but that it has an hereditary sense of direction, or an hereditary impulse to travel in a certain direction, is quite another matter. The sea, to the young bird, may be a barrier ”

“ We do not know the extent of Greenland as a summer breeding home of birds; the growing knowledge of this vast continent proves that its summer avifauna is much larger than we thought, and that western and eastern forms inhabit adjacent breeding areas; the possibility of birds banding with the wrong set of travellers is greater than was suspected.
It is urged that the western shores of Scotland and Ireland should receive these stragglers, but that the records of American birds are fewer from these coasts than from the eastern shores and even Heligoland. ”

“ He never saw a bird at the windows when the moon was shining, and on clear nights the passing crowds go on without a pause. But on hazy nights, with an easterly wind and drizzle, or during fogs, if large numbers of migrants are passing, hundreds may be seen flying in all directions, "all seemingly of the opinion that the only way of escape out of the confusion—is through the windows of the lantern." On one September night, when he was standing on the balcony, he likens the appearance of the birds to a heavy fall of snow. ”

“ Great weight has been placed upon the use of land-bridges and the hereditary habit of crossing seas where these land-bridges once existed but have been submerged during the great geological changes in the earth's surface. Many have insisted that wherever migrants cross the see they do so along submerged coastlines or over submerged land-bridges, arguing that the gradual evolution which has made the advantageous adoption of a habit of migration possible was unable to eliminate the hereditary tendency to follow the exact route by which their ancestors passed from place to place. ”

“ The memory of the bird, call it instinctive memory if we like, would limit the wanderings in search of food to a certain number of places where food was most abundantly found, and the passage between feeding area and breeding area become regular journeys, at the seasons of the year when an increasing number of young birds in the breeding area drove the overgrown population to seek food further from the base, and again when the sexual impulses urged the birds to seek secure nesting sites. ”

“ Some take for granted that the Polar Regions were the original home, the centre of dispersal, of all northern birds, and consequently that migration originated in the gradual pushing back of avian life as the ice gained more and more land each year. During the summer, the birds, urged by an irresistible love of home, travelled as far north as the ice allowed them, but gradually they were driven to nest further and further south until they found refuge in the unglaciated parts of the earth. ”

“ But putting aside economic and utilitarian considerations, there is to some of us a greater stimulus to solve the problems of nature. With the birds, [130] and the insects and plants upon which they feed, we share a common heritage, and the more we learn of the life of these, our fellow workers, the nearer we approach solution of the great riddle of the Universe, the mysterious law-abiding scheme of Nature. The book of knowledge to which we may add some iota is marred with mystery, superstition and error, but each proved fact cleanses its pages. ”

“ In truth there is much to learn, much to which we must still answer—we do not know; but the speculative theory of yesterday is now either myth or fact, and the theory of to-day may be proved true and add something to the data of which knowledge is built. The wildest speculations, based on slender locally ascertained facts or on no foundation whatever except the fertility of the brain, have been offered as solutions of the mysteries; the literature of migration is a jumble of contradictions. ”

“ The solution of the problem is in the interest of science, and therefore of intellectual progress, teaching us the great part which migratory birds play in the scheme of nature. The millions of birds which wander, season after season, from one zone to another, represent an enormous aggregate of labour, by flight and search for food, acting on "the organic life of nature as does the regulator of a steam-engine, at one time accelerating, at another retarding." ”

“ It has been suggested that some of the sexually mature non-breeders may be actually enjoying their winter during our summer; in other words that they have bred in southern breeding-stations whilst their congeners wintered in the same zone. This means a double breeding-area for certain species—a possible explanation, but one hardly supported by known facts. When a bird had so cosmopolitan a range that in the course of its dispersal its breeding areas were separated, we almost invariably find that the birds inhabiting these two areas are distinguishable geographical forms or sub-species. ”

“ From what little we do know about the behaviour of our summer birds in their winter home, we may safely conclude that their habits are similar to those of winter visitors to Britain. Only in a few species are there two restricted areas, two abiding places or homes. The necessity of retaining a secure home for the young and the care of these young during their more helpless age keeps the individual birds within a certain area during the breeding season, but at all other times the bird is more or less of a wanderer. ”

“ Early fall migrants, and indeed the majority of autumn migrants all the world over, travel more slowly than in spring; they are neither impelled by sex-impulses nor the need to escape from failing food supplies. A little later the supply does slacken and with it the temperature cools, and if the changes are sudden southward migration is accelerated. Migration, however, is such an advantageous and well-established habit that it usually begins before hurry is necessary, and the birds loiter southward, feeding as they go. ”

“ That there is an overruling relation between the advance of spring and the passage to northern breeding quarters, and the gradual cooling in autumn and the retreat to winter quarters is, of course, evident, but it must not be held, as contended by the early students of migration, that this is the sole factor which regulates migration. The actual relationship between the weather and the movement of birds is far more complicated than one would imagine, and the stimuli of continental or overland travelling differ from those of a cross-sea flight. ”

“ Possibly in the early days of migration only the mature birds did return; that we cannot state one way or the other. But it is reasonable to argue that once a regular migration habit has become not only confirmed by heredity but a very true advantage to the species, its influence will be felt by each and every individual. Again it is clear that the sexual impulses, in an undeveloped form, are appreciated by the adolescent, and in many animals by even the most juvenile. ”

“ Before the winter resident golden plovers have arrived in autumn and after they have departed in spring, the favourite fields are regularly visited by passing flocks, and the lower reaches of the Mersey, where the common sandpiper is rare as a summer resident, are visited every autumn by parties of birds on passage. Chance may lead a casual wanderer to a good food-supplying spot, but the regularity of appearance suggests habit and memory. ”

“ Birds of Passage or Spring and Autumn Migrants: birds which neither nest with us nor normally remain for the winter, but merely use the British Islands as feeding and resting places on their journey between the northern breeding area and the southern or eastern winter quarters. This group is an especially difficult one, for in it must be included such birds as dunlins and curlews, which are represented as breeding species in Britain, and also a number of birds which apparently go no further south than our islands in winter, and others which, though not breeding, go no further north in summer. ”

“ The natural answer, the cravings of nature and sexual impulses fails to give satisfaction in every case. Wanderings in search of food might lead in any direction, and probably did in the first place, but now birds in the main travel south in search of food and north in search of home, and many of them perform immense journeys, passing over or through lands which are capable of supporting a wealth of bird-life even in the winter months. ”

“ I have seen bands trailing south in autumn in which all the birds were immature, and others in which a number of young were accompanied by a few mature birds, though, certainly, these old birds did not appear to lead. Another assertion, that the old and young do not even travel by the same route rather supports the idea that the young birds find their way simply by a sense of direction, a sense liable to blunder, whilst the old birds travel [61] by the perfected or best route which their experience has taught them. ”