K. Rangachari

“ Some grasses are annual while others are perennial. It is often difficult to determine whether a certain grass is annual or perennial. But by examining the shoot-system this can be ascertained easily. In an annual all the stems and branches usually end in inflorescences and they will all be of the same year. If, on the other hand, both young leafy branches and old branches ending in inflorescences are found mixed, it must be a perennial grass. The presence of the remains of old leaves, underground stolons and rhizomes is also evidence showing the perennial character of the plant. ”

“ So far as the broad general features are concerned, the stems of many grasses are more or less similar in structure. However, when we take into consideration the arrangement of bundles, the development and arrangement of sclerenchyma, every species of grass has its own special characteristics. And these are so striking and constant that it may be possible to identify the species from these characters alone.
We may take as a type the stem of Rottboellia exaltata. ”

“ The first glume is a minute scale. The second glume is shorter than the third and narrower, 5-nerved, ciliate, acute or sometimes with two fine teeth. The third glume is oblong-lanceolate, acute, 5-nerved (rarely 3-nerved) , ciliate on the nerves. The fourth glume is lanceolate, acute, sub-chartaceous, paleate; palea is like the glume in texture. Anthers are yellow and stigmas are white. Lodicules are two and small.
This seems to be a good fodder grass. It grows in all kinds of soils. It is not so common in the plains as on the hills, though it occurs in the plains at the base of the hills. ”

“ The roots are all thin and fibrous in the vast majority of these plants, and they are tough and wiry only in a few cases such as in the case of the roots of Pennisetum cenchroides, P. Alopecuros, Ischæmum pilosum and Andropogon Schœnanthus.
On a close examination it will become evident that all the roots of a grass plant are adventitious. Inasmuch as the growth of the primary root is soon overtaken by other roots growing from the stem, all the roots happen to be of the same size. Roots arise from the nodes just above the insertion of the leaf, and they grow piercing the leaf-sheath. ”

“ The fourth glume and its palea adhere together by their margins. Inside the fourth glume and between it and the palea there are three stamens and an ovary with two styles ending in feathery stigmas. Just in front of the ovary and outside the stamens two very small scale-like bodies are found. These are the lodicules. They are fleshy and well developed in flowers that are about to open. In the spikelet there is only one full flower. The third glume contains no flower in it, but occasionally there may be in its axil three stamens. ”

“ In the case of most grasses the motor-cells are found in groups of three, four or five between the vascular bundles. The central motor-cell is usually the largest and it is somewhat obovate in shape in a transverse section of the leaf. In the leaves of Panicum javanicum and Eriochloa polystachya there are three or four motor cells in the group and the group consists of four, five or rarely six motor cells in the leaves of Eragrostis Willdenoviana. When there are distinct furrows between ridges these cells lie in the furrows and they are many in number. ”

“ And yet very few people appreciate the worth of grasses. Although several families of plants supply the wants of man, the grass family exceeds all the others in the amount and the value of its products. The grasses growing in pasture land and the cereals grown all over the world are of more value to man and his domestic animals than all the other plants taken together.
To the popular mind grasses are only herbaceous plants with narrow leaves such as the hariali, ginger grass and the kolakattai grass. But in the grass family or Gramineæ the cereals, sugarcane and bamboos are also included. ”

“ For example Tragus racemosus grows with all its branches quite prostrate in a poor, dry, open soil. If, on the other hand, this happens to grow in rich soils, or amidst other plants or grasses, it assumes an erect, somewhat tufted habit. Andropogon contortus and Andropogon pertusus are other grasses with a tendency for variation in habit. Plants that are usually small often attain large dimensions under favourable conditions of growth. Ordinarily the grass Panicum javanicum grows only to 1 or 2 feet. ”

“ Instead of being thin and herbaceous it becomes rigid and hard, smooth or rugose externally as in Panicum. Flowering glumes instead of being like empty glumes, become very thin, shorter and hyaline in Andropogon. Sometimes the flowering glumes are awned. All of them may be awned as in Chloris or only the fourth glume as in Andropogon.
The palea is fairly uniform in its structure in many grasses, but it is also subject to variation. It becomes shorter in some and is absent in others. Instead of having two nerves, it may have one and rarely more than two. ”

“ Spikelets are unilateral, sessile, crowded, biseriate on a slender rachis with four to six glumes and 1 to 3-flowered; the rachilla is produced and disarticulating above the empty glumes. The first two glumes are unequal, narrow, keeled, membranous, 1-nerved, persistent, acute, mucronate and the second glume awned shortly. Floral glumes narrow or broad, acute, obtuse or minutely 2-toothed and awned, paleate; sterile glumes are small, without palea. There are two lodicules and anthers are rather small. Grain is narrow and free. ”

“ The leaf-sheaths are shorter than the internodes, flat, compressed, glabrous, with a few hairs or not at the mouth and with membranous margins; the uppermost sheath is spathiform enclosing the inflorescence when young. The ligule consists of only a thin ridge of short hairs densely arranged. Nodes are glabrous and dark-ringed, and with fan-like spreading equitant leaf-sheaths and leaves more especially when rooting. ”