Rachis

Definition and stakes

Portrait of John MacGillivray John MacGillivray,  Narrative of the Voyage of H.M.S. Rattlesnake…

“ Colour of rachis and pinnules, delicate yellowish white above; of rachis, light brown, inferiorly; polypidom about two inches high, rising in several straight simply pinnulated fronds from a common centre; pinnules ascending about 1/4 inch.
4. P. aurita, n. sp.
Cells cup-shaped, tapering at bottom, constricted just below the top; mouth at an angle of 45 degrees, circular; margin subcrenate, plicate, with three folds on each side, with a wide shallow notch in front and entire behind. Rostrum, slender, attenuated below, adnate up to the cell, summit contracted, tubular
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Source: Gutenberg

Portrait of A. C. Seward A. C. Seward,  Fossil plants, Vol. 2: A text-book for students of botany and geology (1910)

“ Below the small-celled epidermis are a few layers of thinner-walled cells which are not arranged in radial series. The structure of the outer part of the cortex is similar to that in the petiole of recent species of Angiopteris (fig. 243, p. 319) and Marattia, in which a more delicate hypoderm is succeeded by a band of mechanical tissue.
The rachis of this type of frond gives off two rows of lateral branches from the vascular axis, the plane of symmetry being at right angles to the primary rachis. Each pinna bore at its base two aphlebiae supplied with vascular strands from the leaf-traces.
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Source: Gutenberg

Marcus Hartog,  The Cambridge natural history, Vol… (1906)

“ The genital rachis itself is an outgrowth from the genital stolon, and this latter originates as a pocket-like ingrowth of the coelom into the wall separating it from the axial sinus; when fully formed it projects into and is apparently contained in this latter space.
Not all the cells forming the genital stolon become sexual cells. Many degenerate and become pigment-cells, a circumstance to which the organ owes its brown colour. In very many species of Starfish many of the cells of the genital rachis undergo a similar degeneration, and hence is produced the apparent aboral blood-vessel.
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Source: Gutenberg

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