Summary

William Swainson Zoological Illustrations, Volume 2

I consider, therefore, formation and sculpture as the only certain characters of species, and that variation of colour should alone distinguish varieties.
It is therefore not surprising that the specific characters given by MM. Bruguiere and Lamarck, and resting principally on the colours of these shells, are frequently obscure, and always long; two inevitable evils attending every attempt to describe minutely the colour, form, and disposition of the markings of shells.
Source: Gutenberg

William Swainson Zoological Illustrations, Volume 2

I have, however, remarked, that the various modifications of the umbilicus, and the termination of the pillar (which is indicated in many species by an elevated ridge or rib within the umbilicus) is a certain and constant indication, presenting the same peculiarity through all the individuals of a species, even in the young state. This termination of the pillar has been mistaken for the inner lip, which, on the contrary, is always above the umbilicus, which, if closed, is not closed by the lip, but by the thickened termination of the pillar or columella.
Source: Gutenberg

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