Various, Encyclopaedia Britannica, 11th Edition…
“ The larvae have the three pairs of legs well developed, and the hinder abdominal segments swollen. Most of the Scarabaeidae are vegetable-feeders, but one section of the family—represented in temperate countries by the dor-beetles (Geotrupes) (fig. 28) and Aphodius, and in warmer regions by the “sacred” beetles of the Egyptians (Scarabaeus) (fig. 27) , and allied genera—feed both in the adult and larval stages, on dung or decaying animal matter. The heavy grubs of Geotrupes, their swollen tail-ends black with the contained food-material, are often dug up in numbers in well-manured fields. ”
