Animal migration

“ In fact, with the entire continent of North America crossed by migratory birds, the different groups or species frequently follow lines that may repeatedly intersect those taken by others of their own kind or by other species. The arterial or trunk routes, therefore, must be considered merely as indicating paths of migration on which the tendency to concentrate is particularly noticeable. ”

“ In fact, bird migration has been described as a phase of geographic distribution wherein there is a more or less regular seasonal shifting of the avian population caused by the same factors that determine the ranges of the sedentary species. If this view is correct, then it must be recognized that the far-reaching works of man in altering the natural condition of the earth's surface can so change the environment necessary for the well being of the birds as to bring about changes in their yearly travels. ”

“ The migration season is full of peril for birds. Untold thousands of smaller migrants are destroyed each year by storms and attacks by predatory animals. These mortality factors, and others, help keep bird populations in check. Perils of migration are among these causes. ”

“ The migration of the survivors, leaving in many cases no representatives behind them, is a difficult problem to solve,—the wingless birds to their special island habitats; the rodents of South America to theirs, leaving the monotremes and marsupials in sole possession of their ancestral domains. ”

“ The relation of the world's land masses to each other and the distribution and association of biotypes within these land masses influence the direction birds migrate. Topography may aid, hinder, or prevent the progress of a migrant depending on the bird's particular requirements. Old World migrants must contend with east-west tending mountain ranges and deserts, whereas New World travelers can proceed north and south across a landscape with its major mountain ranges and river systems oriented in the same direction as the birds migrate. ”

“ Migration owes its origin to the potentiality of flight, enabling birds to advantage themselves by extended dispersals, which through heredity become instinctive, regular and periodical. Geological changes, especially the passing away of the glacial epoch, only influenced by opening up new lands for summer colonisation, but climatic conditions prevented these lands from becoming permanent abodes and fostered the habit of periodical migration. ”

“ Probably northern migrants that winter in equatorial regions and beyond have their migratory urges controlled by similar rhythms or biological clocks. Also, no evidence suggests that the southward migration of birds is controlled by changing periods of light even among species such as white-crowned sparrows, for which this is a controlling factor in the spring. ”

“ Why should a bird subject itself to the rigors of a long migratory journey twice a year if it can find all the requirements suitable for existence in one locality? It seems well to consider briefly the ends that are served by this annual round trip between breeding grounds and winter quarters. Obviously, the migratory habit enables a species to enjoy the summer of northern latitudes and to avoid the severity of winter. In other words, migration makes it possible for some species to inhabit two different areas during the seasons when each presents favorable conditions. ”

“ As we have seen from the foregoing discussion, migration exists in many forms throughout the world and probably arose to satisfy many different needs in different orders of birds at the same time. New pattens, traditions, and routes are arising today as well as disappearing. Currently, the migration patterns we see are a composite result of historic influences mixed with present day influences. Even though the migration of several different species may be very similar, the patterns exhibited today can be the result of quite different evolutionary processes. ”

“ Traditions that have lived for countless generations may die overnight if experienced individuals are lost or no longer active (Hochbaum 1955) ; migration patterns remain constant only as long as the factors influencing these - 97 - patterns remain constant. But the landscape and the interacting ecological stresses are forever changing, and we would expect the adaptive behavior of birds to respond with them. One of these responses to an expanding habitat is colonization of new territory and expansion of a species' range with accompanying development of a migratory habit. ”

“ Birds breeding in the tropics are always resident, except when they breed on mountains, where the climate causes them to descend into the valleys for the winter.”
In many countries we find that while there are some species, such as in England the swallow or the fieldfare, of which every individual disappears at one period of the year or another, there are other species, such as the pied-wagtail or the woodcock, of which only the majority of individuals vanish—a few being always present—and these species form the so-called “partial migrants.” ”

“ In fact, the choice of migration highways is so wide that it seems as if the routes of no two species coincide. Differences in distance traveled, in time of starting, in speed of flight, in geographical position, in latitudes of breeding and of wintering grounds, and in other factors, all contribute to this great variation of migration routes. Nevertheless, there are certain factors that serve to guide the avian travelers along more or less definite lines, and it is possible to define general lines of migration for the majority of species. ”

“ Today we realize that birds do not migrate by "hitching" rides with other birds and that the scope of the migration phenomenon is worldwide, not simply limited to the United States, the Northern Hemisphere, or the world's land masses. The migration heritage is developed just as extensively in Old World warblers migrating to and - 6 - from Europe and Africa as in our wood warblers traveling from Canada and the United States to South America and back. ”

“ Indeed, migration from one region to another has played a most important part in bringing about the present distribution of living things. From what has already been learned as to the past life of the various continents and their shifting connections with one another, it is now feasible to analyze the mammalian faunas of most of them and to separate the indigenous from the immigrant elements. Among the latter may be distinguished those forms which are the much modified descendants of ancient migrants from those which arrived at a much later date and have undergone but little change. ”

“ It affects different birds in different ways. In North America, migratory movements are continent wide, and no evidence has indicated any particular part of the landscape influences all birds in the same manner. Certain bird populations may use general areas in migration, but they are usually not rigidly restricted to them because of topography. ”

“ For more than half a century the Fish and Wildlife Service and its predecessor, the Biological Survey, have been collecting data on the interesting and important phenomenon of the migration of North American birds. The field men of the Service have gathered information concerning the distribution and seasonal movements of the -3- different species in many extended areas, from the Arctic coast south to the pampas of Argentina. ”

“ There probably is no single aspect of the entire subject of bird migration that increases our admiration so much as the unerring certainty with which birds cover thousands of miles of land and water to come to rest in exactly the same spot where they spent the previous summer or winter. Records from birds marked with numbered bands offer abundant proof that the same individuals of many species will return again and again to identical nesting or winter feeding sites. ”

“ To increase such support, information must be more generally available on that little understood but universally fascinating subject of bird migration. A brief presentation of facts on the migratory habits of the birds scientifically gathered by the Fish and Wildlife Service over many years, will be helpful to bird-study classes, to conservation organizations, and to farmers and others individually interested in the welfare of the birds. ”

“ Want of food, and perhaps of the special, proper kind during the breeding season, seems to be the most obvious cause of migration, and none can wonder that those animals which possess the power of removing themselves from a place of scarcity should avail themselves of it, while it is unquestionable that birds possess this faculty in the greatest degree. Even among those species which we commonly speak of as sedentary it is only the adults which maintain their ground throughout the year. ”

“ When in the early days of migration this periodically-supplied northern zoological vacuum was filled to overflowing by the increased numbers of avian inhabitants at the close of the breeding season, the natural food supply would be taxed to its limits; the falling temperature drove some and finally all to seek food further south, and their short migration to lands already filled with old and young birds, caused pressure and overcrowding further south. ”

“ I doubt not that, throughout the ages, geological changes have been an important factor in directing or limiting the scope of migration, and moreover are so still; just as climatic changes and the relative abundance or scarcity of enemies have influenced the course of its evolution. These are all contributory factors operating in the external environment. But there are, besides, internal factors which form part of the inherited constitution of the bird, and, being passed on from generation to generation, afford the conditions under which migration is constantly being renewed. ”

“ Now, in studying the composition of a fauna, and especially its origin, it is of the utmost importance to be able to determine approximately the percentage of accidental arrivals and of the ordinary migrants—that is to say, of those which have reached the country owing to accidental distribution, and of animals which have adopted the usual course of migration. ”

“ While it is beyond question that certain general directions of flight are constantly followed by migratory birds, it is well to remember that the term "migration route" is to some extent a theoretical concept, concerned entirely with the lines of general advance or retreat of a species, rather than the exact course followed by individual birds. ”

“ Much has been learned, and present knowledge is being placed on record, but it must be left to future study to clear away many of the uncertainties that continue to make bird migration one of the most fascinating subjects in the science of ornithology.
Each kind of bird seems to have its own reaction to its environment, so that the character of movement differs widely in the various species, and seldom do any two present the same picture. ”

“ Like the bird's other habits its migratory behavior is just as characteristic as the color of its plumage and, like it, evolved through natural selection because it was advantageous for the survival of the population. Its origin has been thought by some to be a mystery locked in past ages, but by study of the history of how birds came to occupy their present ranges, information becomes available which suggests theories that may be developed and explored. ”

“ We are sometimes told that we must seek the origin of migration in the physical changes that have occurred in the ancient history of the earth—in glacial conditions which gradually forced birds to the south, or in the "stability of the water and mobility of the land" which brought about a gradual separation of the feeding area from the breeding area—and which continued for a sufficient length of time to lead to the formation of an instinct, and that the instinct persists because it is serviceable in promoting the welfare of the race. ”

“ These and other questions on the migratory habits of most species of Northern Hemisphere birds puzzle all who are interested in them, whether it be the farmer who profits by their tireless warfare against the weed and insect pests of his crops, the bird student who enjoys an abundance and variety of feathered inhabitants about him, or the hunter who wants a continuation from year to year of the sport of wildfowling. ”

“ The general anatomical and physiological attributes of birds enable them to develop more diverse and spectacular migratory behavior than any other group of animals. Their potential for long sustained flights is of primary importance in pre-adapting birds to successful migrations. Migration has long since become a definite hereditary habit of many species of birds that recurs in annual cycles, evidently because of physiological changes which prompt a search for an environment suitable for reproduction and survival. ”

“ Typical migration consists of definite movements that are repeated regularly year after year, and it is to these that the term is generally restricted. It is desirable, however, if only for purposes of comparison, that some account be taken of the movements of some other birds, which, while not typical, do possess some of the characteristics of true migration. Data on this subject are being collected through bird banding. ”

“ This suggests that if we are correct in assuming that birds first appeared in a tropical climate, and from such climate migration started, it is probable that by gradually prolonging their northern visits and accustoming themselves to northern insect and vegetable life, these regions became populated by their resident species. It is evident that the present migratory species are simply compelled to return, and three compelling causes are demonstrable. ”