James Bell Pettigrew,
Animal Locomotion; or, walking…
“ The muscles of the wing are so arranged that they can propel it in a horizontal, vertical, or oblique direction. It is a matter of the utmost importance that the direction of the stroke and the nature of the angles made by the surface of the wing during its vibration with the horizon be distinctly understood; as it is on these that all flying creatures depend when they seek to elude the upward resistance of the air, and secure a maximum of elevating and propelling power with a minimum of slip. ”
