Breeding ground

“ But the case would be quite different if the winning sire and dam came from the same old established herd in which the animals had been bred for generations on similar lines. It is this concentration of certain qualities in generation after generation which renders the pedigree animal so intensely prepotent, particularly when mated with animals of an ordinary character or not possessing concentrated breeding. ”

“ Breeders have carried this principle so far as frequently to advocate the growing of plants in the environment most likely to produce the change desired, as, for instance, cultivating tall plants like twining beans in the north or at high altitudes if it is desired to produce a dwarf type or, vice versa, breeding the plants in the south and at a low altitude if a giant or tall type is desired. ”

“ Hence this difference in habituation may well account for the different action of the reproductive organs when domestic races and when species are crossed. It is a nearly analogous fact, that most domestic races may be suddenly transported from one climate to another, or be placed under widely different conditions, and yet retain their fertility unimpaired; whilst a multitude of species subjected to lesser changes are rendered incapable of breeding. ”

“ It is well known that too close breeding, or consanguineous breeding, or breeding in and in, as it is variously called, if continued long, has a bad effect on the offspring, weakening the stock, while judicious crossing of varieties within certain limits of difference has a good effect, strengthening the stock and increasing its fertility. ”

“ The art of breeding has now made such progress, that man can often at discretion produce certain peculiarities in cultivated species 155 of animals and plants. To practised gardeners and farmers, you may give distinct commissions, and say, for example, I wish to have this species of plant with this or that colour, and with this or that shape. Where breeding has reached the perfection which it has attained in England, gardeners and farmers are frequently able to furnish to order the desired result within a definite period, that is, at the end of a number of generations. ”

“ The breeder who wishes to combine fancy and utility properties in any kind of live stock must breed only from specimens that are meritorious in both directions, selecting much more carefully than when breeding for one class of properties. ”

“ If he wish to perpetuate the difference, to form a breed with the peculiarity in question strongly marked, he selects such male and female individuals as exhibit the desired character, and breeds from them. Their offspring are then carefully examined, and those which exhibit the peculiarity the most distinctly are selected for breeding; and this operation is repeated until the desired amount of divergence from the primitive stock is reached. ”

“ From a species in this condition modern poultry breeders have made hundreds of distinct varieties. The easiest method of making a variety in domestication is to select specimens for breeding as near the desired type as possible, and to breed only from a few individuals in each generation which come nearest to the ideal type. ”

“ Suppose then the offspring of any parent to vary in many directions. By cross-breeding among themselves and with the parent stock, these are usually merged in a common type, their differences pooled, and the species remains fixed or else advances slowly by natural selection, along one line, as physical conditions change in geological time. But from time to time there arises a variation in the reproductive organs of some individuals, of such kind that these individuals are fertile among themselves, but sterile or less fertile with other varieties and with the parent stock. ”

“ It is not surprising, seeing the great difference between many domestic breeds, that some few naturalists have concluded that each is descended from a distinct aboriginal stock, more especially as the principle of selection has been ignored, and the high antiquity of man, as a breeder of animals, has only recently become known. Most naturalists, however, freely admit that our various breeds, however dissimilar, are descended from a single stock, although they do not know much about the art of breeding, cannot show the connecting links, nor say where and when the breeds arose. ”

“ These great difficulties in the way of selecting extremes for vegetative propagation are of course met with everywhere. They impede the work of the breeder to such a degree, that but few men are able to surmount them. Breeding new varieties necessitates the bending of every effort to this purpose, and a clear conception of [757] the manifold aspects of this intricate problem. These fall under two heads, the exigencies of practice, and the physiologic laws of variability. ”

“ This now familiar term, proposed by Darwin, was suggested by the operations of breeders in the development and fixation of races for man's use or fancy—breeding in each generation those individuals only in which the desired points are apparent and predominant: in the seed-bed, by rigidly destroying all plants which do not show some desirable variation, breeding in and in from these with strict selection of the most variant form in the particular line or lines, until it becomes fixed by heredity, and as different from the primal stock as the conditions of the case allow. ”

“ Practically all our views on the subject of "races" and their inter-breeding are derived from our observation of the immense number and range of "races" and "breeds" produced by man—as farmer, fancier, and horticulturist. It has been generally received as a rule, that the various races produced in the farm or garden by breeding from a species, will inter-breed freely, and produce offspring which are fertile. ”

“ But for breeding they are found utterly useless; for though they may themselves be uniform in character, they yield during many generations astonishingly diversified offspring. The breeder is driven to despair, and concludes that he will never form an intermediate race. But from the cases already given, and from others which have been recorded, it appears that patience alone is necessary; as Mr. Spooner remarks, “nature opposes no barrier to successful admixture; in the course of time, by the aid of selection and careful weeding, it is practicable to establish a new breed.” ”

“ It is interesting at this point to recall one of the most common differences between plant and animal breeding which is seldom clearly recognized by practical breeders. Plant breeders most commonly strive to produce new races or breeds with distinctive characters which will reproduce their desirable qualities without continuous selection; while animal breeders almost wholly limit their attention to selection within the breeds already established, to maintain them in the highest state of efficiency possible. ”

“ High breeding. When uniformity of plumage can be effected in mixed breeds or varieties without a resort to in-and-in, or close breeding, and without sacrificing the health and vigor of the race, it is desirable ”

“ The disadvantages which the late arrivals have to face are therefore great, and it is probable that the percentage which attain to reproduction will on the average be somewhat lower than the percentage in the case of the earlier arrivals. The district in which my observations have been made lies well within the limits of the breeding range of most of our common species, and it is not surprising that I should have met with little evidence of failure to breed as a result of failure to secure territory. ”

“ St Hilaire great distinction of tame and domestic,—elephants,—ferrets{72}. Reproductive organs not subject to disease in Zoological Garden. Dissection and microscope show that hybrid is in exactly same condition as another animal in the intervals of breeding season, or those animals which taken wild and not bred in domesticity, remain without breeding their whole lives. It should be observed that so far from domesticity being unfavourable in itself «it» makes more fertile: [when animal is domesticated and breeds, productive power increased from more food and selection of fertile races] . ”

“ Both extremes seem to be barred in nature—namely, the inter-breeding of stocks so diverse in structure and quality as to be what we call "distinct species," and again the inter-breeding of individuals of the same immediate parentage or near cousinship. What seems to be favoured by the natural structure and qualities of the plant or the animal is that it shall only breed within a certain group—the species—and shall within that group avoid constant self-fertilization or fertilization by near cousins. ”

“ First, in regard to artificial breeding, we start from the fact last discussed above, viz., that its products in some cases differ from one another much more than the productions of natural breeding. It is a fact that races or varieties often differ from one another in a much greater degree and in much more important qualities than many so-called species, or “good species,”—nay, sometimes even more than so-called “good genera” in their natural state. ”

“ However extensive the varieties produced by artificial breeding, the essential characters of the species remain, and even its minor characters may be reproduced, while the barriers established in nature between species by the laws of their reproduction, seem to be absolute. ”

“ We do not obtain such distributions in describing breeding activity, so we may say that the oscillators regulating such activity are not coupled. Present distribution, habitat preference, residency status, foraging adaptation, previous zoogeographic history, and relicts of ancestral adaptation, all bear on the character of the breeding schedule of any bird species. ”

“ Line-breeding is a systematic form of breeding which preserves the excellence and characteristics of the parent stock. In other words, it perpetuates the blood of both sire and dam so that it is not eventually lost as is the case in cross-breeding. ”

“ These are the two fundamental qualities of animals and plants of which the breeder must avail himself in order to produce new forms. The theoretical principle of breeding is, indeed, extremely simple, but in detail the practical application of this simple principle is difficult and immensely complicated. A thoughtful breeder, acting according to a definite plan, must understand the art of correctly estimating, in every case, the general interaction between the two fundamental qualities of heirship and mutability. ”

“ The breeder is thus enabled through selecting the traits or characters that are wanted and crossing them with a well-known stock, to produce almost any trait or quality that he desires. This law makes it possible to estimate the results of cross breeding with almost mathematical exactness. Improved varieties of fruits, grains and vegetables have been produced in this manner, and with animals marvelous results have been achieved. ”

“ The risk attending too close breeding as in the breeding of pure breds is also avoided provided that the herd from which the sires are bought is sufficiently large to furnish a change of blood, yet of similar breeding. ”

“ Conspicuous breeding activity occurs from the first third of May to the last third of June.
The Influence of Dominant Foraging Adaptation.—Five categories here considered reflect broad foraging adaptation: woodland species, taking invertebrate foods in the breeding season from woody vegetation or the soil within wooded habitats; grassland species, taking invertebrate foods in the breeding season from within grassland situations; limnic species, foraging within marshy or aquatic habitats; aerial species, foraging on aerial arthropods; raptors, feeding on vertebrates or large insects. ”

“ The rapid development of the science and art of breeding places us to-day in position to secure improvements much more rapidly than has been done in the past. It would not be astonishing if from 25 to 50 years of careful, intelligent breeding would accomplish with a wild plant what has required many centuries under the crude methods of our ancestors.
It may be asked why we should be in haste to take up the improvement of our native plants. ”

“ With our true domestic animals, on the other hand, fertility is perfect, and is very little affected by changed conditions. Thus, we see the common fowl, a native of tropical India, living and multiplying in almost every part of the world; and the same is the case with our cattle, sheep, and goats, our dogs and horses, and especially with domestic pigeons. It therefore seems probable, that this facility for breeding under changed conditions was an original property of the species which man has domesticated—a property which, more than any other, enabled him to domesticate them. ”

“ It follows from this that the capacity of breeding at an early age and at short intervals, as with pigeons, rabbits, etc., facilitates selection; for the result is thus soon made visible, and perseverance in the work encouraged. It can hardly be an accident that the great majority of the culinary and agricultural plants which have yielded numerous races are annuals or biennials, which therefore are capable of rapid propagation, and thus of improvement. ”

“ The slight differences, however, which characterise each individual animal and plant would in most, probably in all cases, suffice for the production of distinct races through careful and prolonged selection. We see what selection, though acting on mere individual differences, can effect when families of cattle, sheep, pigeons, &c., of the same race, have been separately bred during a number of years by different men without any wish on their part to modify the breed. ”