The Journal of Indian Botany (1919)
“ Spurs with more or less than three leaves are very common in seedlings. Eifty-seven per cent, seedlings possesses such abnormal spurs. 3. Spurs with 4-leaves are the commonest of all, being 83'8 % of all abnormal shoots. 4. The conclusion is drawn from these facts that a 3-leaved spur has been derived from a spur with more leaves, that the spur is equi- valent to an ordinary shoot and that pines with a small number of needles in their spurs are more highly specialised than species with a larger number of needles. ”
