Various, Encyclopaedia Britannica, 11th Edition…
“ Haeckel considers that the Siphonophores have two distinct ancestral lines of evolution:1. In the Disconanthae, i.e. in such forms as Velella, Porpita, &c., the ancestor was an eight-rayed medusa (Disconula) which acquired a pneumatophore as an ectodermal pit on the ex-umbrella, and in which the organs (manubrium, tentacles, &c.) became secondarily multiplied, just as they do in Gastroblasta as the result of incomplete fission. The nearest living allies of the ancestral Disconula are to be sought in the Pectyllidae. ”
