E. S. Russell, Form and Function: A Contribution to the History of Animal Morphology
“ The stapes forms a close connection with the hammer and the incus. In birds, where there is a single ear-ossicle, the columella, the middle piece of arch I forms, as we have seen, the quadrate, by means of which the lower jaw is joined to the skull. The proximal end of Meckel's cartilage, which in mammals forms the hammer, here gives the articular surface between the lower jaw and the quadrate. The columella is formed from the middle piece of the three into which the cartilage of the second arch segments. It is, therefore, the homologue of the stapes in mammals. ”
