Charles Darwin,
The Various Contrivances by which Orchids are Fertilised by Insects
(1877)
“ In some of the cases above specified in which the pollinia apparently undergo no movement of depression, I am not sure that there was not a very slight one after a time. In the various Ophreæ the anther-cells are sometimes seated exteriorly and sometimes interiorly with respect to the stigma; and there are corresponding outward and inward movements in the pollinia: but in the Vandeæ the anther-cells always lie, as far as I have seen, directly over the stigma, and the movement of the pollinium is always directly downwards. ”
