Equidae

“ Gaudry places the Quagga nearest of all living Equidae to the Hipparion gracile of Pikermi.
Fossil Equidae.—The existing Equidae all belong to the genus Equus, though there are some who would (quite unnecessarily) divide off the Zebras as a genus Hippotigris. The genus Equus itself goes back in time to the Pliocene, during which epoch there lived in India E. sivalensis, the same species according to some with the E. stenonis of Europe. None of these species, Old World or New, are easily to be separated from E. caballus. ”

“ No hair-clad mammal either within the family of the Equidæ, or without, has any other arrangement on the under surface of its neck than what is here shown to be the normal one—a uniform uninterrupted slope from the head to the chest. There is also a feature of this greatly variegated piece of the horse’s coat under its neck, and that is that it is so highly variegated with diversity of pattern as to make it unlike any normal or natural structure or character in any animal. That is not the way Nature does her normal work. ”

“ A very easy answer to this is found by studying, not only any Ungulate known, except the Gnu, but more particularly all wild Equidæ; and this reveals the fact that in all this series the normal slope of hair prevails here, that is to say, an even trend from head to shoulder. Variations in others, indeed, hardly exist, and I may add that the absence of variations here is a strong piece of negative evidence in my favour, for no Ungulate comes near the domestic horse for amount and activity of locomo­tion, which is indeed his raison d’être. He is the only one that has invented new patterns. ”

“ If a horse of the hackney type and a zebra were skinned and the bodies of the two animals then examined I suppose a competent anatomist would find some difficulty in distinguishing one from the other so closely do these two allied species of equidæ, one wild and the other domesticated, resemble one another in structure. But in this as in many other questions form is not to be considered alone. The coloura­tion of the two animals is strikingly different, but, in its humble way, the difference of their patterns of hair-arrangement is worthy of notice. ”

“ While, therefore, the ancient progenitors of the Equidæ were common to North America and Europe, in Miocene and even Eocene times, true horses appear to have arisen in the Palæarctic region, to have passed into North America in the latter part of the Pliocene period, and thence to have spread over all suitable districts in South America. They were not, however, able to maintain themselves permanently in their new territory, and all became extinct; while in their birth-place, the Old World, they continue to exist under several varied forms. ”

“ The development of the limbs of the Horse shows a most interesting series, of stages, which correspond in part to the ancestral forms which palaeontology seems to prove to be the line of the descent of our existing Equidae. ”

“ As you know, there was no trace of the genus horse on our island continent before the coming of the white man. In America, on the other hand, although there was no horse as we know him, before the advent of the Conqueror Cortez, in 1518, yet the fossilised remains of the Eohippus, the Protohippus and Hipparion are so numerous and well distributed on the great American continents that these wide lands seem to have been the most favoured home of the great race of equidae, in the far-off days before the ice. ”

“ The zebras (3 species) inhabit the greater part of the Ethiopian region, while the asses (4 species) are characteristic of the deserts of the Palæarctic region from North Africa and Syria to Western India, Mongolia, and Manchuria. The domestic horse is not known in a wild state, but its remains are found in recent deposits from Britain to the Altai Mountains, so that its disappearance is probably due to human agency.
Extinct Equidæ.—Extinct forms of this family are very numerous. The genus Equus occurs in Post-pliocene and Pliocene deposits in Europe, North America, and South America. ”

“ The dentition is generally the full one of forty-four teeth. The premolars in the more ancient forms are unlike the molars, but like them in more recent forms. The molars of the upper jaw have two crests parallel and united by an outer crest. The fore-feet have four, the hind-feet three toes.
The family is fully as ancient as that of the Equidae, but the specialisation of the toes never advances so far. The modern representatives of the order are, so far as the feet are concerned, in the condition of very early representatives of the equine stock. ”

“ It is hence not surprising that many are led to inquire to which particular branch of the equine family tree the Shetland pony belongs.
Not many years ago it was said the horse tribe was at the start represented by a primitive race about the size of a fox but with as many hoofs as a tapir—four in front and three [150] behind,—that in course of time a new race arose (with three hoofs in front as well as behind) which eventually gave rise to the one-toed ancestor of all the living Equidæ. ”

“ The molar teeth are hypselodont, and the premolars, with the exception of the first, resemble the molars in their pattern. The orbit is completely surrounded by bone. The incisors are chisel-shaped, with a pit on the free surface. The canines are rudimentary if present. The radius and ulna are fused, as are the tibia and fibula. Although for the sake of uniformity a family, Equidae, is here separated from its allies, it is quite impossible owing to the full state of our knowledge of this group to draw a really hard-and-fast line between this family ”

“ As certain individuals are grouped by naturalists into species, so particular species, which in habits and general appearance resemble each other, are arranged under the head of genus. The horse, the ass, and the zebra are formed on nearly the same anatomical plan; they are therefore classed together, and designated the genus Equus, a term derived from the Latin word equus, a horse—that animal being regarded as the type, or perfect member of the group. Thus the horse, in the nomenclature of the naturalist, is termed Equus caballus ”

“ A good case by which to exhibit further our conception of a species is that afforded by the species which are united in the genus Equus—the horse-genus. There are living at the present day several wild kinds of Equus—namely, the wild horse, or Tarpan, of the Gobi desert of Mongolia, called after the Russian explorer Przewalski; two kinds of Asiatic wild ass, called the Kiang and the Onegar; the African wild ass, and two or three kinds of zebra. There are, besides, many kinds of domesticated horses, ranging from the Shetland pony to the Flemish dray horse, and from the Shire horse to the Arab. ”

“ Beginning with the Perissodactyla, or odd-toed ungulates, we find the Equine animals remarkably numerous and interesting. The true horses of the genus Equus, so abundant in the Post-Pliocene formations, are represented in the Pliocene by several ancestral forms. The most nearly allied to Equus is Pliohippus, consisting of animals about the size of an ass, with the lateral toes not externally developed, but with some differences of dentition. ”

“ But the appearance of the stripes on the various breeds of the horse, when of a dun-colour, does not afford nearly such good evidence of their descent from a single primitive stock as in the case of the pigeon; because no certainly wild horse is known as a standard of comparison; because the stripes when they do appear are variable in character; because there is far from sufficient evidence of the appearance of the stripes from the crossing of distinct breeds; and lastly, because all the species of the genus Equus have the spinal stripe, and several have shoulder and leg stripes. ”

“ He had a foot forming almost a straight line with the leg, which, however, did not prevent it from being turned in, so that it was an equinus together with something of a varus, or else a slight varus with a strong tendency to equinus. But with this equinus, wide in foot like a horse’s hoof, with rugose skin, dry tendons, and large toes, on which the black nails looked as if made of iron, the clubfoot ran about like a deer from morn till night. He was constantly to be seen on the Place, jumping round the carts, thrusting his limping foot forwards. ”

“ We might have presumed that as no living representative of the equine family, whether horse, ass, zebra, or quagga, had been furnished by North or South America when those regions were first explored by Europeans, a search in the transatlantic world for fossil species might be dispensed with. ”

“ The true Horses are represented by one well-established species, Equus caballus, from which all the other races, or varieties, are descended, by a process of selection under the care of man, and these vary in size, proportion of parts, and colour, as much as any two closely-allied species of wild animals can be said to be defined from each other. According to Mr. Darwin, no aboriginal or truly Wild Horse is positively known to exist, for the Wild Horses of the East may probably be descended from those which have escaped from the service of man. ”

“ The only advantage these large mares may give to the mule is in the size of the feet and bone that they may impart. The heavier you can get the bone and feet, the better. And yet you can rarely get even this, and for the reason that I have before given, that the mare, in nineteen cases out of twenty, breeds close after the jack, more especially in the feet and legs. It makes little difference how you cross mares and jacks, the result is almost certain to be a horse's body, a jack's legs and feet, a jack's ears, and, in most cases, a jack's marks. ”

“ These leg bars were at birth more distinct than in the Zebra sire. Continuous with the mane is a well-defined dorsal band (with a narrow yellow band at each side) which extends some distance into the tail. The tail in the hybrid had, at birth, long hairs right up to the root, but, notwithstanding this, there were three distinct bars visible at each side; similar tail bars I have once seen in a Horse.
Though the ears look long in some of the photographs, they are now relatively very little longer (though rounder at the apex) than in the majority of Horses. ”

“ Now, as to the living forms or form-kinds of the genus Equus—which are we to regard as true species, and which are only varieties and races of lower significance than species? The answer is clear enough in regard to several of them. The wild Mongolian horse and all the domesticated horses are varieties, races, or breeds of one species, judged not only by such marks as the possession of callosities on both the hind and the fore legs, but also by the test of breeding. They breed together and produce persisting races. ”

“ There were, as it happened, only three pinto horses in the entire saddle stock, and these three were the last left of the entire bunch. Now a little boy or girl, and many an older person, thinks that a spotted horse is the real thing, but practical cattle men know that this freak of color in range-bred horses is the result of in-and-in breeding, with consequent physical and mental deterioration. It was my good fortune that morning to get a good mount of horses,—three sorrels, two grays, two coyotes, a black, a brown, and a grulla. ”

“ The selection of a mare, relatively larger than the horse, is an important rule in breeding, and it is believed that much of the success of Arabian and other Eastern horses as stock-getters, has resulted from the application of this principle. They possess valuable traits, but condensed within too small a compass. When such an animal is put to a well-bred, larger mare, the fœtus has abundance of room and nourishment to develop and perfect the circumscribed outlines of the male parent, and acquire for itself increased volume and character. ”

“ All the existing members of the family, such as the domesticated horse (Equus caballus) and its wild or half-wild relatives, the asses and the zebras, are included in the typical genus. In all these the crowns of the cheek-teeth are very tall (fig. 1, b) and only develop roots late in life; while their grinding-surfaces (fig. 2, b and c) are very complicated and have all the hollows filled with cement. The summits of the incisors are infolded, producing, when partially worn, the “mark.” In the skull the orbit is surrounded by bone, and there is no distinct depression in front of the same. ”

“ As a substitute for turning horses out for a brief run in the spring, it is customary in some stables to cut grass and give it, varied by vetches and clover, to the horse in his box. These salutary alteratives are good in themselves, but there is still wanting the glorious fresh air of the open paddock, which, when all nature is awakening from the long slumber of winter, is so renovating to the equine system.
It is best to fetch your horse up at night, because it is in the night when turned out that he eats the most ”

“ With three manes, equine, awful, majestic, down to their sides. Fierce heroes who wield against foeman hard-smiting swords. A blow, they give with three iron flails having seven chains triple-twisted, three-edged, with seven iron knobs at the end of every chain: each of them as heavy as an ingot of ten smeltings. Three big brown men. Dark equine back-manes on them, which reach their two heels. Two good thirds of an oxhide in the girdle round each one's waist, and each quadrangular clasp that closes it as thick as a man's thigh. ”

“ The remains of equine mammals, now known from the tertiary and quaternary deposits of this country, already represent more than double the number of genera and species hitherto found in the strata of the eastern hemisphere, and hence afford most important aid in tracing out the genealogy of the horses still existing. ”