Various, McClure's Magazine, Vol. 1, No…
“ All the essential parts of the middle ear—the tympanic membrane, tympanic cavity, and Eustachian tube—develop from the first gill-opening with its surrounding parts, which in the Primitive Fishes (Selachii) remains throughout life as an open blowhole, situated between the first and second gill-arches. In the embryos of higher Vertebrates it closes in the centre, the point of concrescence forming the tympanic membrane. The remaining outer part of the first gill-opening is the rudiment of the outer ear-canal. ”
