Herbert Spencer,
The Principles of Biology, Volume 2…
(1899)
“ That is to say, while the power of flight is retained at all, the muscles and bones cannot greatly dwindle; but the dwindling, in birds whose flights are short or infrequent or both, will be in the reserve stock of the substance that is incapacitated by action, or in the appliances that keep the apparatus in repair, or in both. Only where, as in the struthious birds, the habit of flight is lost, can we expect atrophy of all the parts concerned in flight ”
